In the following passage, some of the words have been left out. Read the passage carefully and select the correct answer for the given blank out of the four alternatives.
In order to reason ____________ it is not merely necessary to have a good intellect. An athlete may have the proper _____________, good framework, and symmetrical muscles, but he cannot expect to cope __________ others of his kind unless he has learned to develop ___________ muscles and to use them to the best advantage. And, in the same way, the man who wishes to reason correctly must develop his intellectual faculties and must also learn the art __________ using them to the best advantage.
Identify the most appropriate option to substitute the underlined segment. If no substitution is required, select No substitution. The shoebill stork can be recognise from its long legs and broad wings.